Phylum: Bryozoa
Class: Stenolaemata
Order: Trepostomatida
Family: Atactotoechidae
Genus: Cyphotrypa Ulrich & Bassler, 1907
Cincinnatian Species: Cyphotrypa clarksvillensis, Cyphotrypa frankfortensis, Cyphotrypa madisonensis, Cyphotrypa osgoodensis, Cyphotrypa waynensis

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Type Species: Leptotrypa acervulosa (Ulrich, 1893)
History: (Nickles & Bassler, 1900)

Species found in the Cincinnatian (

  • Cyphotrypa clarksvillensis (McFarlan, 1931)
  • Cyphotrypa madisonensis (Brown & Daly, 1985)
  • Cyphotrypa pachymuralis (Brown, 1965)
  • Cyphotrypa switzeriensis (Karklins, 1984)
  • Cyphotrypa waynensis (Utgaard & Perry, 1964)
  • Cyphotrypa osgoodensis (Perry and Hattin, 1960)
  • Cyphotrypa frankfortensis (Ulrich & Bassler)

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Stratigraphic Occurrences

Geologic Range
Ordovician – Devonian

Geographic Occurrences

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Sequences (Formations)

  • C5 Sequence (Lower Whitewater, Waynesville: Blanchester, Clarksville, Ft. Ancient)
  • C4 Sequence (Arnheim)
  • C2 Sequence (Fairview: Fairmount)
  • C1 Sequence (Clays Ferry/Kope: Economy/Fulton; Lexington/Pt. Pleasant)

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Identification in Hand Sample:

  • Zoarium Morphology: Massive
  • Zoecia: Polygonal; thin-walled; acanthopores well developed
  • Mesozooids: None
  • Monticules: Low
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Steven Holland (UGA Strat Lab, 2013)

  • Cyphotrypa is massive, rarely ramose, with low monticules. Zooecia are polygonal and thin-walled, with diaphragms common in exozone. Lacks mesozooids. Styles have large clear lumen, and are typically common and occur at junctions of walls.

Karklins(1984 p. I-45)

  • Remarks: The original definition of the genus Cyphotrypa, by Ulrich and Bassler (1904, p. 29), is followed here. Ulrich and Bassler (1904) characterized Cyphotrypa as having massive zoaria in which autozooecia have thin walls, polygonal shapes in cross section, and abundant basal diaphragms and styles (acanthopore of Ulrich and Bassler), but lack mesozooecia (their mesopores). C. switzeriensis n. sp. described here, conforms to this characterization except for its apparent absence of styles in zoaria.

Treatise on Invertebrate Paleontology, Part G (1953):

  • Massive, attaining width of 25 cm. Zooecia thin-walled, with diaphragms in recurrent mature zones. Mesopores absent; acanthopores well developed.

McFarlan (1931)

  • Massive heterotrypids with thin-walled, prismatic zooecia. Diaphragms and acanthopores well developed. Mesopores wanting

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C. clarksvillensis

C. frankfortensis