Phylum: Bryozoa
Class: Stenolaemata
Order:  Cryptostomata
Genus: Escharopora Hall, 1847
Cincinnatian Species: Escharopora falciformis, Escharopora hilli, Escharopora maculata, Escharopora pavonia

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Type Species: Escharopora recta (Hall, 1847)
Species found in the Cincinnatian, USA

  • Escharopora acuminata (James)
  • Escharopora eparmata (Karklins, 1983)
  • Escharopora falciformis (Nicholson)
  • Escharopora hilli (Ulrich)
  • Escharopora maculata (Ulrich)
  • Escharopora pavonia (D’Orbigny)


    Note: PBDB notes the order of this genus as Fenestrata. However, the Invertebrate Treatise places it within the order Cryptostomata.

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Geographic Occurrences

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Sequences (Formations)

  • C3 Sequence (Mt. Auburn, Corryville)
  • C2 Sequence (Fairmount, Mount Hope)
  • C1 Sequence (McMicken, Southgate, Economy, Fulton)

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Identification in Hand Sample:

  • Zoarium Morphology:Unbranching, simply “frond”
  • Zooecia: Sub-elliptical; Arranged diagonally in intersecting series (angles between 50* and 80*)
  • Mesozooids: Few
  • Monticules: Absent to common; flat to slightly raised; may be irregular or form transverse ridges across frond.
  • Maculae:

Species Differentiation

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Steve Holland (2013, UGA strat lab):

  • Unbranched and lanceolate zooarium (bifoliate and proximally tapering); rarely branched. Subelliptical zooecia aligned or alternating across zooarium

Treatise on Invertebrate Paleontology. Part G (1983):

  • Zoarium generally unbranched and lanceolate, rarely branched; connecting segments with tapered proximal tips, probably articulating with encrusting bases. Mesothecae straight to sinuous; autozooecial ranges aligned or alternating across mesotheca. In exozones, autozooecia form angles between 50* and 80* with mesothecae, subelliptical in cross section. Autozooecial wall and stereom laminae form sinuous striae at zoarial surface. Pustules common along autozooecial boundaries, striae scattered in exozonal walls and stereom. Living chambers subelliptical in cross section. Superior hemisepta common, generally blunt and short, rarely thin and long, curving proximally, usually scattered in zoaria, but may be regularly arranged. Mural spines absent to common, irregularly shaped, scattered in zoaria; may be regularly arranged. Exilazooecia few, subelliptical to subcircular in cross section, commonly closed at zoarial surfaces by thickened walls, sparse in zoarial margins and in proximal zoarial parts, generally absent in zoarial mid regions. Monticules absent to common, flat to slightly raised, irregularly shaped or may form annular ridges at regular intervals across zoaria.

McFarlan (1931):

  • Zoarium a simple or branching frond, articulating with a basal expansion. Zooecial apertures in diagonally intersecting series, surrounded by sloping hexagonal areas.

Bassler (1911):

  • Although long considered a synonym of Ptilodictya, the species of Escharopora form a natural assemblage related to the former genus in its general zoarial and zooecial characters but differing mainly in the arrangments of their apertures. In Ptilodictya the zooecial apertures are arranged in regular longitudinal rows; in Escharopora their arrangement is in diagonally intersecting series. The internal structure of the two genera is essentially the same.
  • All of the species of Escharopora are of Ordovician age while the oldest known forms of Ptilodictya occur in the earliest Silurian (Richmond) deposits. The close relationship between the two genera is thus apparent from their geologic distribution, Ptilodictya being undoubtedly a derivative of the earlier genus.
  • Genotype.—Escharopora recta Hall. Middle Ordovician of New York and Canada.

Nickles & Bassler (1900):

  • Like Ptilodictya, but distinguished chiefly by the arrangement of the apertures, which are in decussating series, and by the sloping hexagonal area which surrounds the apertures.
  • Genotype: Escharopora recta Hall. Ordovician. Fourteen described, and eleven new species.

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E. falciformis

E. hilli

E. maculata

E. pavonia